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Title: The Origin of Species
Description: reproductive isolation, biological species concept, speciation,
Description: reproductive isolation, biological species concept, speciation,
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Chapter 24: The Origin of Species
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• such barriers block gene flow between species limit the forma>on of hybrids, offspring that result
from an interspecific ma>ng
• Prezygo'c barriers (before the zygote) block fer>liza>on from occurring
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by preven>ng an aCempted ma>ng from being completed successfully
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2 Specia6on can take place with or without geographic sepera6on
• allopatric specia'on, gene flow is interrupted when a popula>on is divided into geographically
isolated subpopula>ons
• ex-‐ water level in a lake may subside, resul>ng in two or more smaller lakes that are now home to
separated popula>ons
The Process of Allopatric Specia6on
• one geographic separa>on has occurred, the separated gene pools may diverge
• different muta>ons arise, the natural selec>on and gene>c driQ may alter allele frequencies in
different ways in the separated popula>ons
Polypoloidy
• a species may originate from an accident during cell division that results in extra sets of chromosomes,
a condi>on called polyploidy
• polyploid specia>on occasionally occurs in animals
• two dis>nct forms of polyploidy have been observed in plant popula>ons
• autopolyploid is an individual that has more than two chromosomes sets that are all derived from a
single species
• ex-‐ a failure of cell division could double a cell’s chromosome number from the original number (2n)
to a tetraploid number (4n)
• a tetraploid offspring can produce fer>le tetraploid offspring by self-‐pollina>ng or by ma>ng with other
tetraploids — reproduc>vely isolated from 2n plants of the original popula>ons, because the triploid
(3n) offspring of such unions have reduced fer>lity
• a second form of polyploidy can occur when two different species interbreed and produce hybrid
offspring
• in subsequent genera>ons, various mechanisms can change a sterile hybrid into a fer>le polyploid
called an allopolyploid, fer>le when ma>ng with each other but cannot interbreed with either parent
species; thus, they represent a new biological species
Habitat Differen6a6on
• sympatric specia>on can occur when a subpopula>on exploits a habitat or resource not used by the
parent popula>on
Title: The Origin of Species
Description: reproductive isolation, biological species concept, speciation,
Description: reproductive isolation, biological species concept, speciation,