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Title: Human evolution
Description: Human evolution

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Human  Evolution  
 

 
The  Oxford  debate  1860  
 
Thomas  Henry  Huxley,  nicknamed  'Darwin's  bulldog',  was  championing  Charles  Darwin's  
revolutionary  concept  of  evolution  by  natural  selection,  published  less  than  a  year  before
...
 Both  sides  claimed  victory,  and  the  debate  
continues  to  this  very  day
...
 Who  
made  so  ill  a  use  of  his  wonderful  speaking  powers  to  try  and  burke,  by  a  display  of  
authority,  a  free  discussion  on  what  was,  or  was  not,  a  matter  of  truth”  
 
Paleoanthropological  story  –  Lucy  
 
During  the  1960s  and  1970s,  hundreds  of  fossils  were  found,  particularly  in  East  
Africa  in  the  regions  of  the  Olduvai  gorge  and  Lake  Turkana
...
 
 
They  found  "Lucy",  the  most  complete  fossil  member  of  the  species  Australopithecus  
afarensis
...
   
 
All  arisen  from  a  single  ancestral  type  that  once  lived  on  a  continent  now  submerged  
beneath  the  Indian  Ocean
...
 Single  dispersal?  
2
...
 Dispersal  from  North  or  South?  
 
OR    a  multiregional  origin  of  modern  humans  which  allows  for  a  much  greater  role  for  
hybridisation
...
   
 

Roughly  100,000  years  ago,  the  Old  World  was  occupied  by  a  morphologically  diverse  group  
of  hominids
...
   
 
However,  by  30,000  years  ago  this  taxonomic  diversity  vanished  and  humans  everywhere  
had  evolved  into  the  anatomically  and  behaviorally  modern  form
...
 
 
Understanding  the  issue  
 
Multiregional  theory:  homo  erectus  left  Africa  2  mya  to  become  homo  sapiens  in  different  
parts  of  the  world
...
   
 
This  model  contains  the  following  components:  
-­‐   Some  level  of  gene  flow  between  geographically  separated  populations  prevented  
speciation,  after  the  dispersal  
-­‐   All  living  humans  derive  from  the  species  Homo  erectus  that  left  Africa  nearly  two  
million-­‐years-­‐ago  
-­‐   Natural  selection  in  regional  populations,  ever  since  their  original  dispersal,  is  
responsible  for  the  regional  variants  (sometimes  called  races)  we  see  today  
-­‐   The  emergence  of  Homo  sapiens  was  not  restricted  to  any  one  area,  but  was  a  
phenomenon  that  occurred  throughout  the  entire  geographic  range  where  humans  
lived  
 
In  contrast,  the  Out  of  Africa  Model  asserts  that  modern  humans  evolved  relatively  recently  
in  Africa,  migrated  into  Eurasia  and  replaced  all  populations  which  had  descended  
from  Homo  erectus
...
 
-­‐   After  Homo  erectus  migrated  out  of  Africa  the  different  populations  became  
reproductively  isolated,  evolving  independently,  and  in  some  cases  like  the  
Neanderthals,  into  separate  species  
-­‐   Homo  sapiens  arose  in  one  place,  probably  Africa  (geographically  this  includes  the  
Middle  East)  
-­‐   Homo  sapiens  ultimately  migrated  out  of  Africa  and  replaced  all  other  human  
populations,  without  interbreeding  

Modern  human  variation  is  a  relatively  recent  phenomenon  
 
The  multiregional  view  posits  that  genes  from  all  human  populations  of  the  Old  World  
flowed  between  different  regions  and  by  mixing  together,  contributed  to  what  we  see  today  
as  fully  modern  humans
...
 As  these  peoples  migrated  they  replaced  all  other  
human  populations  with  little  or  no  interbreeding
...
 
 
 
 
 
 
 
 
What’s  wrong  with  mitochondrial  Eve?  
•   Africa  has  greater  genetic  diversity  because  its  prehistoric  population  was  probably  
larger    
•   Assumes  genetic  diversity  reflects  age  of  a  population  rather  than  population  size    
•   Differences  in  ancient  population  size  could  mimic  a  recent  African  origin  of  modern  
humans
Title: Human evolution
Description: Human evolution